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Cagalli and her golden mobile suit Akatsuki have turned the tide of battle in Orb's favor, but she now finds herself facing enemy reinforcements, in the form of the Minerva and Shinn's Destiny Gundam. Even as the Archangel rushes to Cagalli's aid, Kira and Lacus return from space with a pair of new Gundams.

Aboard the Archangel, the injured Athrun thinks back upon the path he has followed to this point. Even though he saw the pointlessness of war during the previous conflict, he rejoined ZAFT and fought alongside the Minerva crew in order to bring the new war to an end.

Shinn and Rey receive medals for their exploits at Heaven's Base, and Chairman Durandal awards them their own FAITH insignia. But a new crisis is already beginning, as ZAFT has discovered that Lord Djibril has taken refuge in Orb.

The Minerva follows the Girty Lue into the debris belt that surrounds Earth, and sends Shinn and his comrade Lunamaria Hawke in pursuit of the enemy mothership. However, Neo has tricked the ZAFT pilots into following a decoy.

ZAFT has captured Heaven's Base only to find that Lord Djibril has fled to parts unknown, and Chairman Durandal orders his forces to track the fugitive down. Aboard the Archangel, Kira laments his own powerlessness and wonders what will happen next.

The ZAFT forces gather around Heaven's Base as they wait for the Earth Forces to respond to Chairman Durandal's ultimatum. Aboard the Minerva, Lunamaria channels her grief into anger at Logos, while Shinn does his best to comfort her.

As Athrun and Meyrin escape from Gibraltar in a stolen mobile suit, Shinn and Rey pursue them in the new Destiny and Legend Gundams. Claiming that the fugitives have stolen classified information, Chairman Durandal orders Shinn and Rey to shoot them down.

Chairman Durandal, accompanied by Meer Campbell, presents Shinn and Athrun with a pair of powerful new Gundams. Shinn is delighted with this gift, but Athrun is still upset that they were ordered to attack the Archangel.

Although the Archangel has escaped the Minerva's attack, the Freedom has been destroyed, and the ship itself badly damaged. After retrieving Kira, the Archangel continues on to Orb, while Shinn returns to the Minerva and receives a hero's welcome.

Chairman Durandal has revealed the existence of Logos to the public, throwing the world into a state of confusion. Kira and Cagalli fear that their homeland is now in danger, and the Archangel begins making its way back to Orb.

After laying Stella's body to rest, Shinn returns to the Minerva with a new determination. When Athrun visits Shinn's room, he finds Shinn and Rey running combat simulations to prepare for a confrontation with the Freedom.

The Destroy Gundam advances towards Berlin, crushing one city after another. The Archangel arrives to confront the Destroy, but even Kira's Freedom Gundam is no match for its overwhelming power.

Shinn and Rey have returned Stella to her allies in order to save her life. As punishment for defying military regulations, the two pilots are placed in detention while the ZAFT leadership determines their fate.

In the aftermath of the battle at Crete, the Archangel retrieves survivors from the Orb fleet, who ask to join Cagalli in her fight to restore Orb's ideals. Meanwhile, the Minerva is undergoing another round of repairs before continuing on to Gibraltar.

Shinn and Rey struggle against the enemy commander Neo Roanoke. In order to help her pilots, ZAFT Captain Talia Gladys orders the newly constructed warship Minerva to launch from Armory One and join the battle. The enemy forces withdraw, and the Minerva gives chase.

Lacus returns to space and joins her allies aboard the warship Eternal, which is now hidden inside a cluster of asteroids. Chairman Durandal orders his forces to continue searching for Lacus, insisting that she is merely an impostor.

As the Minerva struggles against the combined Orb and Earth Forces fleets, the Freedom Gundam and the Archangel appear on the battlefield once more. Cagalli pleads with the Orb forces to cease fighting, but Yuna orders them to continue their assault on the Minerva.

Even as Neo prepares for another attack on the Minerva, the ZAFT forces are on the trail of the Archangel and its crew. While Chairman Durandal supervises the pursuit of the fugitive Lacus, Captain Talia of the Minerva receives a report on Athrun's secret meeting with Kira and Cagalli.

After running off alone to the Lodonia laboratory, the Extended soldier Stella Loussier has been captured by the Minerva's pilots. Shinn brings the injured Stella aboard the Minerva, but the ship's doctors are unable to treat her due to her abnormal physiology.

Athrun has arranged a meeting with Kira and Cagalli, but the old friends no longer see eye to eye. Even after hearing of the assassination attempt on Lacus, Athrun still believes in Chairman Durandal's good intentions, and he returns to the Minerva after urging his friends to keep Orb from participating in the war.